A pet chameleon (Johnny Depp) becomes accidentally stranded in the Mojave Desert after his terrarium falls from his owner's car. After meeting an armadillo named Roadkill (Alfred Molina), who is seeking the mystical Spirit of the West, he narrowly avoids being eaten by a red-tailed hawk. The next day, after having a surreal nightmare, he meets desert iguana Beans (Isla Fisher), a rancher's daughter, who takes him to Dirt, an Old West town populated by desert animals.
Beans discovers that the water reserves, stored in a water-cooler bottle in the bank, are dangerously low. At the Gas Can Saloon, the chameleon, using bravado and improvisation to fit in, presents himself as Rango, a tough drifter. He quickly runs afoul of outlaw Bad Bill (Ray Winstone), narrowly avoiding a shootout when the hawk returns, scaring Bill. The hawk chases Rango until by luck Rango kills the predator by crushing it under an empty water tower that he accidentally caused to collapse. In response, the Mayor (Ned Beatty) appoints Rango the new sheriff. A skeptical Beans demands Rango investigate the water problem while the townsfolk worry that the hawk was the only thing keeping gunslinger Rattlesnake Jake from returning to terrorize them.
Ashamed and no longer knowing who he is, Rango wanders the desert and, in a daze, meets the Spirit of the West (Timothy Olyphant), a cowboy whom Rango calls the Man with No Name. The Spirit inspires Rango and tells him, "No man can walk out on his own story". With the aid of Roadkill and mystical moving cacti, Rango learns the source of Dirt's water is Las Vegas and that someone has shut off a water line. Realizing the mayor's hand in this, Rango recruits the hill clan in his plan.
Returning to town, he calls out Jake for a duel—a diversion so that the hill folk and the cacti can flood the town with water. The mayor threatens Beans' life, forcing Rango to surrender. The two are put into the bank vault to drown, while the mayor prepares to shoot Jake, whom he calls a relic. However, Rango manages to take the only bullet from the gun and uses it to break the door of the vault, flooding the room and taking out the mayor and his men. Jake, acknowledging Rango as a worthy opponent and a great legend for saving his life, grabs the mayor and drags him into the desert to take revenge. The citizens of Dirt celebrate the return of the water and acknowledge Rango as their hero.
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